Hoot Omega

Hoot Omega is the rampaging avian Guardian who serves as the boss of Highvault. Upon his defeat, he drops for Nina, or  for Ace.

Attacks
Hoot Omega does not attack much himself; he will initially start the fight by shooting a spread of three lightning bolts at the player, then flying away. The player must pursue him through various platforming puzzles, while under siege from a constant rain of lightning bolts. Occasionally, Hoot Omega will summon three tornadoes close together, indicated by a column of red ! marks that indicate where the tornadoes are going to appear.

At the end of each chunk, you will find Hoot Omega waiting for you and he will fight you directly, occasionally launching projectiles at you while you attempt to fight him. Once significantly damaged (about a third of his HP taken away), Hoot Omega will fly away once more and the player must repeat this cycle. Also, until he is forced to fly away, a much larger tornado will linger behind him, barring the player's progress until Hoot Omega is forced to run away or until the player waits enough time, after which it will disappear.

After about three chunks, Hoot Omega will be cornered in one last room, and will not run away anymore, permitting the player to attack him all they like.

Tips
Nina's (dropped from Echobeast) will temporarily freeze Hoot Omega; not only that, but freezing him will stop the thunder rain for the rest of the fight.

Trivia
Hoot Omega is, amusingly, the only boss that does not explode on death; instead he will simply fall to the ground facefirst.